PhD in Biochemistry, University of Cambridge
- Principal Scientist, Imperial Cancer Research Fund (ICRF) Clare Hall Laboratories (- present)
- Cancer Research UK (until December 2011)
| 1991 | Fellow of the Royal Society (-present) |
| 1999 | Foreign Associate of the US National Academy of Sciences (- present) |
| 2005 | Member, Board of Governors, Okinawa Institute of Science and Technology Promotion Corporation (until October 2011) |
| 2006 | Chairman of EMBO Council (until 2010) |
| September 2009 | Establishing Member of Okinawa Institute of Science and Technology School Corporation (until October 2011) |
| November 2011 | Member, Board of Governors, Okinawa Institute of Science and Technology School Corporation (-present) |
| 1990 | Nobel Prize in Medicine |
