| 1974 |
Ph.D. in Chemistry, Harvard University |
| 1979-1985 |
Assistant Professor of Neurobiology, Harvard Medical School |
| 1985-1995 |
Associate Professor of Neurobiology, Harvard Medical School |
| 1991-1993 |
Director, Graduate Program in Neurosciences, Harvard University |
| 1996-1999 |
Director, Division of Fundamental Neuroscience, National Institute of Neurological Disorders and Stroke (NINDS) |
| 1999-2007 |
Associate Director for Technology Development, Office of the Director, NINDS |
| 2005-2007 |
Special Adviser, OIST |
| 2007(January) |
Senior Adviser to the President, OIST |
| 2007(September) |
Vice President and Executive Director, OIST |
